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Planta (2005), in press.

The Agrobacterium rhizogenes rolC-gene induced somatic embryogenesis and shoot organogenesis in Panax ginseng transformed calluses

Gorpenchenko 1 T.Y., Kiselev 1 K.V., Bulgakov 1 V.P., Tchernoded 1 G.K., Bragina 2 E., Khodakovskaya 1 M.V., Koren 1 O.G., Batygina 2 T.B., Zhuravlev 1 Yu.N.

1- Institute of Biology and Soil Sciences, Far East Division, Russian Academy of Sciences, pr. Stoletiya Vladivostoka 159, Vladivostok, 690022 Russia; fax: 7 (4232) 31-0193; e-mail: bulgakov@ibss.dvo.ru 2- Komarov Botanical Institute of Russian Academy of Sciences, 2 Prof. Popov Street, St.-Petersburg, 197376, Russia

Abstract
Expression of the Agrobacterium rhizogenes rolC gene in Panax ginseng callus cells results in formation of tumors that are capable to form roots. The selection of the tumor clusters, which did not develop roots, yielded the embryogenic 2c3 callus line. The ability to form somatic embryos is independent of external growth factors. Such developmental shift is of particular interest because this is the first indication of the rolC-embryogenic effect and, to our knowledge, the first indication that a single gene of non-plant origin can induce somatic embryogenesis in plants. The DNA sequence analysis showed that the somatic-to-embryonic transition was not due to a mutation of the rolC gene inserted. Although the 2c3 somatic embryos developed through a typical embryogenesis process, they terminated prematurely and repeatedly formed adventitious shoot meristems and embryo-like structures. A part of the shoots and somatic embryos formed enlarged and fasciated meristems. The observed developmental abnormalities are similar to those that have been previously described for Arabidopsis embryogenic calluses where the WUSCHEL gene was continuously overexpressed.